Clausiliidae of mainland China (Gastropoda, Stylommatophora)

Hartmut Nordsieck

General remarks

The clausiliid fauna of mainland China is very rich. In a revision of the Chinese Clausiliidae (Nordsieck 2001, 2002), the first since the basic work of Gredler, Heude, Moellendorff and O. Boettger hundred years ago, the number of species amounted to 138. Most species (134) were classified with the subfamily Phaedusinae, the remaining four with the subfamily Garnieriinae. In the few years since then the species number has increased to 185 (169 Phaedusinae species, 16 Garnieriinae species). The new species have already been described (Nordsieck 2003, 2005, 2007b) or are being prepared for description. The increase in this short span shows that the majority of Chinese species is still unknown.
If one compares the clausiliid fauna of mainland China with that of all Asia, about one third of all Phaedusinae species and nearly half of all Garnieriinae species have been found in this country. Therefore, concerning clausiliid diversity, mainland China is one of the most interesting areas of Asia. One has to consider, however, that the knowledge of the other countries of Asia diverges a lot; the well-known Japanese fauna contrasts with that of major parts of Southeast Asia which are nearly unexplored.
This discrepancy in knowledge is also true of the clausiliid faunae within mainland China itself. In this country Clausiliidae occur only south of a border-line which can be drawn from northern Sichuan along Shaanxi and Henan to Jiangsu. From this area only the faunae of the provinces Guangxi, Hubei and Sichuan (with Chongqing) are comparatively well-known. In contrast, the knowledge of the faunae of Yunnan, Hunan, Jiangxi, Anhui and Zhejiang is insufficient, while those of the remaining provinces are nearly unknown.
A determination of Chinese clausiliid species is difficult, because there is no useful reference work for identification. In principle, the shell is sufficient for determination, not only because the shell morphology is characteristic for each species, but also because of only a few species the anatomy, especially the genital morphology, is known (Nordsieck 2007a: 67). The aim of this work is to give a survey on the Chinese clausiliids by the characteristics of their genera and a list of their species, with information on their distribution. Collectors and other interested persons are thus enabled to determine at least the genus a species belongs to. Subsequently, they can try to identify the species by studying the respective literature and reliable voucher material of species of that genus kept in the collection of a museum.
If using shell morphology for determination, one should know how species are distinguished, i. e. the characters of taxonomic importance. As is also the case with other clausiliids, a determination of Chinese clausiliids is not possible without studying their closing apparatus which is only present in adult specimens. Its structure and the respective terms are presented in Nordsieck 2007a (: 177-178). In Chinese clausiliids especially the inferior lamella, the palatal plicae and the clausilium plate are taxonomically important.

Systematic list
(Provinces with type locality in bold)

The two subfamilies occurring in mainland China, Phaedusinae and Garnieriinae, are easily distinguished by the characters of the aperture (Nordsieck 2007a: 11, 13). In the following the genera of both subfamilies are characterized by shell morphology, and the species, with information on their distribution, are listed. A list of synonyms is added for each subfamily.

Phaedusinae A. J. Wagner 1922

Synprosphymini H. Nordsieck 2007

Synprosphyma A. J. Wagner 1920

Synprosphyma (Synprosphyma)
Yunnan, Sichuan (+ Chongqing), Guangxi, Hunan, Hubei, Shaanxi.
basilissa (Boettger & Schmacker 1894) (b. basilissa, b. planicollis H. Nordsieck 2007): Sichuan.
cavicola (Gredler 1888): Hubei.
cervicalis (Bavay & Dautzenberg 1909): Yunnan.
cookei (Preston 1912): Shaanxi.
duplex (H. Nordsieck 2003): Yunnan.
ehrmanni (H. Nordsieck 2001): Sichuan.
fistulata (Bavay & Dautzenberg 1909): Yunnan, Guangxi.
franciscana (Moellendorff 1885) (f. dayongensis (Chen & Zhang 1999), f. franciscana): Sichuan (Chongqing), Hubei, Hunan.
fraterminor (Gredler 1892): Hubei.
gibbosula (Deshayes 1870): Sichuan.
imperatrix (Boettger & Schmacker 1894): Sichuan.
lyra (Gredler 1887): Hunan.
n. sp. 1: Sichuan.
n. sp. 2: Sichuan.
n. sp. 3: Sichuan.
n. sp. 4: Sichuan.
succinea (Heude 1886): Yunnan.
thibetiana (Deshayes 1870): Sichuan.
vulpina (Heude 1886): Sichuan (Chongqing), Hubei.
wanshiensis (Blume 1927): Sichuan (Chongqing).
xichangensis (Chen & Zhang 1999): Sichuan.
yanghaoi Grego & Szekeres 2011: Guangxi.

Synprosphyma (Excussispira) Lindholm 1925
Sichuan (Chongqing), Hubei.
decurtata (Heude 1885) (d. decurtata, d. leiostoma (Heude 1890)): Sichuan (Chongqing).
fargesiana (Heude 1885): Sichuan (Chongqing).
fargesianella (Heude 1885): Sichuan (Chongqing).
inversa (Heude 1886): Sichuan (Chongqing).
laurentiana (Moellendorff 1885): Hubei.
lunatica (Heude 1888): Sichuan (Chongqing).
pseudinversa (H. Nordsieck 2001): Sichuan (Chongqing).
retorta (Heude 1890): Sichuan (Chongqing).

Phaedusini

Oospira Blanford 1872

O. (Oospira)
Yunnan, Guangxi, Guizhou, Guangdong.
?baziniana (Heude 1889): Guangxi.
?binaria (Heude 1886): Yunnan.
gastroptychia (Moellendorff 1885): Guizhou.
gerlachi (Moellendorff 1881): Guangdong.
siderea (Heude 1886): Yunnan.

O. (Siphonophaedusa) Lindholm 1924
Guangxi.
grangeri (Bavay & Dautzenberg 1899) (g. asiphonia H. Nordsieck 2007, g. grangeri): Guangxi.

O. (Atractophaedusa) Ehrmann 1927
Guangxi.
ookuboi H. Nordsieck 2005: Guangxi.
takagii H. Nordsieck 2005: Guangxi.
zhaoyifani Grego & Szekeres 2008: Guangxi.

O. (Distortiphaedusa) Grego & Szekeres 2011
Guangxi.
imprimata (Grego & Szekeres 2011): Guangxi.

O. (Formosana) O. Boettger 1877
Yunnan, Sichuan (+ Chongqing), Guangxi, Guizhou, Guangdong, Hunan, Hubei, Jiangxi, Anhui, Zhejiang.
antilopina (Heude 1885) (a. antilopina, a. unicornu (Boettger & Schmacker 1894)): Sichuan (Chongqing), Hubei.
artifina (Heude 1885): Sichuan (Chongqing), Hubei, Guizhou.
dextrogyra (Bavay & Dautzenberg 1909): Guizhou, Guangxi, Guangdong.
friniana (Heude 1890): Anhui, Zhejiang.
kiangshiensis (Gredler 1892) (k.kiangshiensis, k. n. subsp. Hunan): Hunan, Jiangxi.
kongshanensis H. Nordsieck 2007: Yunnan.
liujinae Maassen 2008: Guangxi.
magnaciana (Heude 1882): Sichuan (Chongqing).
moschina (Gredler 1888): Hubei.
moschinella H. Nordsieck 2007: Guizhou.
n. sp.: Guangxi.
n. sp.: Sichuan.
ooharai H. Nordsieck 2007: Yunnan.
pacifica (Gredler 1884) (p. n. subsp. Anhui, p. n. subsp. Hunan, p. pacifica, p. siantanensis (Gredler 1885)): Hunan, Anhui.
psilodonta (Heude 1889): Hubei.
schawalleri H. Nordsieck 2001: Sichuan.
semprinii (Gredler 1884): Hunan.
splendens H. Nordsieck 2005 (s. n. subsp. Hunan, s. splendens): Guizhou, Hunan, Guangxi.
telum H. Nordsieck 2003: Sichuan (Chongqing).
umbrosa H. Nordsieck 2007: Guizhou.

O. (Paraformosana) H. Nordsieck 2003
Sichuan (Chongqing), Hubei.
delavayana (Heude 1885): Sichuan (Chongqing).
indurata (Heude 1886): Sichuan (Chongqing), Hubei.
jinyungensis (Chen & Zhang 1999): Sichuan (Chongqing).
lepidospira (Heude 1889): Hubei.
longispina (Heude 1885): Sichuan (Chongqing).
phyllostoma (Heude 1888): Sichuan (Chongqing), Hubei.
plicilabris (Grego & Szekeres 2011): Sichuan (Chongqing).

O. (Formosanella) H. Nordsieck 2003
Yunnan, Sichuan (+ Chongqing), Hubei.
bensoni (H. Adams 1870): Sichuan (+Chongqing), Hubei.
recedens (Boettger & Schmacker 1894) (r. n. subsp. Sichuan, r. recedens): Sichuan (+ Chongqing), Yunnan.
?seguiniana (Heude 1885): Yunnan.

Dautzenbergiella Lindholm 1924

D. (Dautzenbergiella)
Guangxi.
chinensis H. Nordsieck 2007: Guangxi.

D. (Mansuyiella) H. Nordsieck 2003
Guangxi.
mansuyi (Dautzenberg & Fischer 1908) (m. mansuyi, m. ootanii H. Nordsieck 2007): Guangxi.

Acanthophaedusa H. Nordsieck 2007
Guangxi.
ookuboi H. Nordsieck 2007: Guangxi.
reductans Grego & Szekeres 2011: Guangxi.

Miraphaedusa H. Nordsieck 2005
Guangxi.

M. (Miraphaedusa)
takagii H. Nordsieck 2005: Guangxi.

M. (Falsiluna) Grego & Szekeres 2011
harryleei (Grego & Szekeres 2011): Guangxi.

Fuchsiana Gredler 1887
Hunan.
paradoxa (Gredler 1883): Hunan.

Bacillophaedusa Grego & Szekeres 2011
Hubei.
annularis (Grego & Szekeres 2011): Hubei.

Serriphaedusa H. Nordsieck 2001
Yunnan, Sichuan.
boisseaui Grego & Szekeres 2011: Sichuan.
?cholerigena (Heude 1889): Hubei.
draesekei (Blume 1925) (d. caviphila H. Nordsieck 2003, d. draesekei): Sichuan.
elamellata (Boettger & Schmacker 1894): Sichuan.
kangdingensis (Chen & Zhang 1999): Sichuan.
marteli (Dautzenberg 1915): Sichuan.
n. sp. 1: Sichuan.
n. sp. 2: Sichuan.
n. sp. 3: Sichuan.
n. sp.: Yunnan.
ookuboi H. Nordsieck 2007: Sichuan.
serrata (Deshayes 1870) (s. n. subsp. Sichuan, s. serrata): Sichuan.
violacea Grego & Szekeres 2011: Sichuan.

Sinigena Lindholm 1925
Sichuan (Chongqing).
bisdelineata (Heude 1885): Sichuan (Chongqing).
vincotiana (Heude 1885): Sichuan (Chongqing).

Macrophaedusa Moellendorff 1883
Jiangxi, Anhui, Zhejiang.
cecillii (L. Pfeiffer 1847): Zhejiang.
frankei (Boettger & Schmacker 1894): Jiangxi.
veruta (Heude 1890) (v. oliveriana (Annandale 1924), v. veruta): Anhui, Zhejiang.

Macrophaedusella H. Nordsieck 2001
Anhui.
jesuitica H. Nordsieck 2001: Anhui

Bathyptychia Lindholm 1925

B. (Bathyptychia)
Yunnan, Sichuan (+ Chongqing), Hubei.
aplostoma (Heude 1885): Sichuan (Chongqing).
beresowskii (Moellendorff 1902) (b. beresowskii, b. delicata H. Nordsieck 2003, b. flavida H. Nordsieck 2005, b. n. subsp. Sichuan): Sichuan, Yunnan.
bulimina (Gredler 1892): Hubei.
hupeana (Gredler 1892): Hubei.
hupecola (Gredler 1888): Hubei.
infantilis (Gredler 1890): Hubei.
martensi H. Nordsieck 2001: Sichuan.
mira H. Nordsieck 2003: Sichuan.
n. sp.: Shaanxi.
n. sp.: Sichuan.
provisoria (Gredler1888): Hubei.

B. (Strictiphaedusa) H. Nordsieck 2001
Jiangxi.
strictilabris (Schmacker & Boettger 1890): Jiangxi.

B. (Brachyptychia) H. Nordsieck 2001
Yunnan, Hubei.
breviplica (Moellendorff 1886): Hubei, Yunnan.
recens (Gredler 1894): Hubei.

Celsiphaedusa H. Nordsieck 2001
Hubei.
celsa (Gredler 1892): Hubei.

Streptodera Lindholm 1925
Hubei.
trachelostropha (Moellendorff 1885): Sichuan (Chongqing), Hubei.

Hemiphaedusa O. Boettger 1877

H. (Selenophaedusa) Lindholm 1924
Yunnan, Guangxi.
bavayi H. Nordsieck 2002: Guangxi.
billeti (Fischer 1898) (b. polydonella H. Nordsieck 2005): Guangxi.
diplochilus (Moellendorff 1901) (d. griffithsi (Grego & Szekeres 2011)): Guangxi.
lantenoisi (Dautzenberg & Fischer 1905): Yunnan.
media H. Nordsieck 2005: Guangxi.
ooharai H. Nordsieck 2007: Guangxi.
spinifera H. Nordsieck 2005: Guangxi.
thatkheana (Bavay & Dautzenberg 1899) (t. ptychostoma H. Nordsieck 2011, t. splendida H. Nordsieck 2011): Guangxi.

H. (Hemiphaedusoides) H. Nordsieck 2001
Jiangxi, Fujian, Anhui, Zhejiang.
moellendorffiana (Heude 1882) (m. edentula (Boettger & Schmacker 1894), m. moellendorffiana): Anhui, Zhejiang.
ringens (Schmacker & Boettger 1890): Jiangxi.
thaleroptyx (Moellendorff 1882): Fujian.

H. (Labyrinthiphaedusa) H. Nordsieck 2001
Zhejiang.
labyrinthoides (Sykes 1898): Zhejiang.

H. (Hemiphaedusa)
Fujian, Zhejiang.
pluviatilis (Benson 1842) (p. pluviatilis, p. zilchi H. Nordsieck 2001): Zhejiang, Fujian.

H. (Margaritiphaedusa) H. Nordsieck 2001
Yunnan, Sichuan, Guangxi, Guizhou, Hunan, Hubei, Jiangxi.
cristina (Gredler 1888): Hubei.
hensaniensis (Gredler 1901): Hunan.
macroptychia H. Nordsieck 2007: Guizhou.
margaritifera (Bavay & Dautzenberg 1909): Yunnan.
n. sp.: Hunan.
principalis (Gredler 1881): Hunan.
protrita (Gredler 1887) (p. hunancola H. Nordsieck 2001, p. protrita): Hunan, Jiangxi.
rusticana H. Nordsieck 1998 (r. amoena H. Nordsieck 2001, r. rusticana): Sichuan, Hubei.
ziyuanensis (Chen & Zhang 1999): Guangxi.

H. (Hemizaptyx) Pilsbry 1905
Guangxi, Guizhou, Hubei, Fujian, Anhui, Jiangsu (+ Shanghai).
heudeana (Moellendorff 1882): Jiangsu, Shanghai, Anhui.
minuta Yen 1939: Hubei, Guizhou, Guangxi, Fujian.

Phaedusa H. & A. Adams 1855

P. (Phaedusa)
Yunnan, Sichuan, Guangxi, Guizhou, Guangdong, Hainan, Hubei.
bocki (Sykes 1895) (b. bocki, b. menglunanensis (Chen & Zhang 1999)): Yunnan.
boettgeri H. Nordsieck 2001: Yunnan.
coccygea (Gredler 1888): Hubei.
elisabethae (Moellendorff 1881) (e. elisabethae, e. matejkoi Grego & Szekeres 2011, e. pingguonensis (Chen & Zhang 1999)): Guangxi, Guangdong.
hainanensis (Moellendorff 1884): Hainan.
lypra (Mabille 1887) (l. debaonensis Chen 1994, l. latestriata H. Nordsieck 2007, l. naponensis (Chen & Zhang 1999)): Guizhou, Guangxi.
potanini Moellendorff 1902 (p. potanini, p. pretiosa H. Nordsieck 2001): Sichuan.
sorella H. Nordsieck 2003: Yunnan, Guizhou.
szechuanensis Pilsbry 1934: Sichuan.

P. (Metaphaedusa) H. Nordsieck 2001
Sichuan, Guangxi, Hunan, Hubei, Shaanxi.
hilberi (O. Boettger 1884): Shaanxi.
n. sp.: Sichuan.
pallidocincta (Moellendorff 1886) (p. ookuboi H. Nordsieck 2005, p. pallidocincta): Hunan, Guangxi.
pseudaculus H. Nordsieck 2001: Hubei.
pseudobensoni (Boettger & Schmacker 1894) (p. aberrans H. Nordsieck 2003, p. pseudobensoni): Sichuan.

Reinia Kobelt 1876

R. (Reinia)
Fujian.
eastlakeana (Moellendorff 1882): Fujian.

Euphaedusa O. Boettger 1877

E. (Dentiphaedusa) H. Nordsieck 2003
Yunnan, Sichuan (+ Chongqing), Guangxi, Guizhou, Hubei, Anhui.
presbyteralis (Gredler 1892): Hubei.
spinula (Heude 1882) (s. clavulus (Heude 1886), s. diaconalis (Heude 1889),
s. spinula): Yunnan, Sichuan (+Chongqing), Guizhou, Guangxi, Hubei, Anhui.
E. (Euphaedusa)
Yunnan, Sichuan (+ Chongqing), Gansu, Guangxi, Guizhou, Guangdong, Hunan, Hubei, Shaanxi, Jiangxi, Fujian, Anhui, Jiangsu (+ Shanghai), Zhejiang.
aculus (Benson 1842) (a. aculus, a. insularis (Heude 1882), a. labio (Gredler 1878), a. nankingensis (Heude 1882)): Zhejiang, Shanghai, Jiangsu, Anhui, Jiangxi, Hubei, Sichuan (+Chongqing), Guizhou, Hunan, Guangxi, Fujian.
anceyi (O. Boettger 1882): Shaanxi, Hubei.
belemnites (Boettger & Schmacker 1894): Hubei.
buccinella (Heude 1886): Sichuan (Chongqing), Guizhou.
costifera H. Nordsieck 2001: Sichuan.
cylindrella (Heude 1886) (c. crobylodes (Schmacker & Boettger 1890), c. cylindrella, c. medioglabra (Schmacker & Boettger 1894), c. n. subsp. Guizhou): Yunnan, Guizhou, Hubei.
fitzgeraldae (O. Boettger 1879): Fujian?
krejcii H. Nordsieck 2001: Sichuan.
loczyi (O. Boettger 1884) (l. acanthula (Heude 1885), l. loczyi, l. pagnucciana (Heude 1888), l. parietaria (Schmacker & Boettger 1890)): Yunnan, Sichuan (+Chongqing), Gansu, Shaanxi, Hubei, Guizhou, Hunan, Jiangxi, Guangxi, Fujian.
lunanensis Chen & Zhang 1999 (l. acutissima H. Nordsieck 2003, l. lunanensis): Yunnan.
minatoi H. Nordsieck 2003: Sichuan.
planostriata (Heude 1882): Anhui, Jiangxi.
ridicula (Gredler 1882): Hunan.
simiola (Gredler 1883): Hunan.
steetzneri (Pilsbry 1919): Sichuan.
yunnancola H. Nordsieck 2001: Yunnan.

E. (Papilliphaedusa) H. Nordsieck 2003
Yunnan, Sichuan (Chongqing), Guangxi, Guizhou, Guangdong (+ Hongkong), Fujian.
constellata (Heude 1886): Sichuan (Chongqing), Guizhou.
kunmingensis Chen & Zhang 1999: Yunnan.
lorraini (Menke 1856): Guangdong (Hongkong).
mucronifera H. Nordsieck 1998: Fujian.
porphyrea (Moellendorff 1882) (p. yangshuoensis (Chen 1994), p. porphyrea, p. theristica (Mabille 1887)): Guangxi, Guangdong.
yongshuae H. Nordsieck 2007: Guizhou.

E. (Tauphaedusa) H. Nordsieck 2003
Guangdong (+ Hongkong), Hunan, Jiangxi, Fujian, Anhui, Jiangsu (+ Shanghai), Zhejiang.
broderseni (Boettger & Schmacker 1894): Guangdong (Hongkong).
gemina (Gredler 1881) (g. davidi H. Nordsieck 2001, g. gemina): Hunan, Jiangxi.
ponsonbyi (O. Boettger 1883): Fujian?
tau (O. Boettger 1877) (t. cetivora (Heude 1882), t. hunana (Gredler 1881), t. janseniana (Heude 1885)): Zhejiang, Shanghai, Anhui, Fujian, Hunan, Guangdong.

E. (Telophaedusa) H. Nordsieck 2003
Sichuan (+ Chongqing), Guizhou, Hubei, Hunan, Shaanxi.
faberi (Boettger & Schmacker 1894): Sichuan.
filippina (Heude 1882): Sichuan (Chongqing), Hubei, Hunan.
gredleri H. Nordsieck 1998: Hubei.
ishibei H. Nordsieck 2007: Guizhou.
latilunellaris (Boettger & Schmacker 1894): Sichuan (Chongqing), Hubei, Guizhou.
lea (Gredler 1892): Hubei.
longurio (Boettger & Schmacker 1894): Guizhou.
meroniana (Heude 1888): Shaanxi.
praecelsa (Gredler 1884): Hubei.
socia (Gredler 1892): Hubei.
tetsui (Boettger & Schmacker 1894): Hubei.

Synonyms
(Emendations and wrong determinations omitted)

adaucta Gredler 1889 [non A. & G. Villa] = Euphaedusa gredleri;
albopapillata Schmacker & Boettger 1890 = Oospira lepidospira;
aprivora Heude 1886 = Synprosphyma franciscana;
beipeinensis Chen & Zhang 1999 = Oospira bensoni;
caudata Gredler 1888 = Phaedusa coccygea;
cecillei Boettger & Schmacker 1894 [non L. Pfeiffer] = Macrophaedusa veruta;
chamelodonta Gredler 1890 = Oospira psilodonta;
changyangensis Chen & Zhang 1999 = Euphaedusa filippina;
chinensis L. Pfeiffer 1849 = Hemiphaedusa pluviatilis;
chinensis Moellendorff 1874 [non L. Pfeiffer] = Euphaedusa aculus;
chongqingensis Chen & Zhang 1999 = Oospira magnaciana;
circinnata Heude 1889 = Phaedusa lypra?;
clarocincta Boettger 1883 = Macrophaedusa cecillii;
coelicola Gredler 1890 = Euphaedusa spinula;
colombeliana Heude 1882 = Euphaedusa aculus;
comminuta Heude 1889 = Euphaedusa porphyrea?;
constricta Heude 1890 = Synprosphyma decurtata;
continua Pilsbry 1934 = Phaedusa pseudobensoni;
crassa Yen 1939 = Oospira indurata;
cyclostoma Moellendorf 1883 [non L. Pfeiffer] = Euphaedusa tau;
cyclostomella Zilch 1951 = Euphaedusa tau;
danbaensis Luo, Chen & Zhang 1998 = Oospira bensoni;
discreta Boettger & Schmacker 1894 = Oospira recedens;
distorta Küster 1861 = Euphaedusa aculus;
emeiensis Luo, Chen & Zhang 1998 = Oospira bensoni;
flavescens Heude 1884 = Euphaedusa tau;
fortunei L. Pfeiffer 1852 = Macrophaedusa cecillii;
frigida Heude 1884 = Euphaedusa aculus;
fuchsiana Heude 1885 = Fuchsiana paradoxa;
fulvella Heude 1882 = Euphaedusa aculus;
gigas Moellendorff 1885 [non O. Boettger] = Oospira indurata;
gracilis Yen 1939 = Euphaedusa anceyi;
guanxianensis Chen & Zhang 1999 = Phaedusa pseudobensoni;
guanyangensis Chen & Zhang 1999 = Hemiphaedusa minuta;
hunkonensis Chen & Zhang 1999 = Euphaedusa planostriata;
infecta Heude 1886 = Streptodera trachelostropha;
jansseni Chen & Zhang 1999 = Euphaedusa spinula;
jiulaodongensis Chen & Zhang 1999 = Oospira bensoni;
julii Gredler 1888 = Phaedusa hilberi;
labrosa Heude 1886 = Oospira indurata;
laevisculpta Yen 1939 = Euphaedusa latilunellaris;
lamothei Ancey 1906 = Oospira siderea;
largillierti L. Pfeiffer 1847 = Hemiphaedusa pluviatilis;
leucospira Heude 1882 = Euphaedusa aculus;
liedtkei Blume 1925 = Synprosphyma thibetiana;
lingyunensis Chen & Zhang 1999 = Euphaedusa spinula;
loloensis Heude 1890 = Bathyptychia breviplica;
longxianensis Chen, Liu & Xu 1996 = Euphaedusa anceyi;
ludingensis Chen & Zhang 1999 = Bathyptychia beresowskii;
magnacianella Heude 1882 = Oospira bensoni;
margaritacea Heude 1886 = Synprosphyma franciscana;
microthyra Schmacker & Boettger 1894 = Euphaedusa spinula;
missionis Ancey 1883 = Euphaedusa aculus;
moellendorffi E. Martens 1875 = Euphaedusa aculus;
mucronata Moellendorff 1882 [non Held] = Euphaedusa mucronifera;
novemgyrata Ancey 1885 = Euphaedusa spinula;
novemspiralis O. Boettger 1884 = Euphaedusa loczyi;
orphanuli Heude 1882 = Euphaedusa aculus;
orthoceras Schmacker & Boettger 1894 = Euphaedusa spinula;
oscariana Gredler 1892 = Bathyptychia breviplica;
pachyodon Heude 1884 = Hemiphaedusa heudeana;
pachystoma Heude 1882 [non L. Pfeiffer] = Hemiphaedusa heudeana;
papillacea Gredler 1884 = Euphaedusa porphyrea;
papillina Gredler 1890 = Oospira lepidospira;
polytropa Schmacker & Boettger 1894 = Euphaedusa spinula;
purpurascens Moellendorff 1885 = Oospira artifina;
rathouisiana Heude 1882 = Euphaedusa tau;
ruptiva Heude 1889 = Phaedusa coccygea;
rustica Heude 1889 [non A. Schmidt] = Hemiphaedusa rusticana;
sanmenensis Luo, Chen & Zhang 1998 = Euphaedusa tau;
sangpingensis Chen & Zhang 1999 = Oospira phyllostoma;
sarcochila Ancey 1883 = Hemiphaedusa heudeana;
scholastica Heude 1889 = Euphaedusa spinula;
septemlamellata Ancey 1883 = Euphaedusa aculus;
septemplicata Heude 1882 = Euphaedusa aculus;
shanghiensis L. Pfeiffer 1853 = Euphaedusa aculus;
siangnanensis Yen 1939 = Euphaedusa anceyi;
straminea Heude 1882 = Euphaedusa tau;
superaddita Heude 1882 = Euphaedusa aculus;
timalthea Sykes 1898 = Oospira friniana;
tumefaciens Chen & Zhang 1999 = Oospira bensoni;
vinacea Heude 1882 = Euphaedusa aculus;
wentschuanensis Blume 1925 = Oospira bensoni;
weigoldi Blume 1927 = Euphaedusa spinula;
yaanensis Chen 1994 = Oospira bensoni;
yunnanensis Sykes 1895 = Phaedusa bocki;
yunyangensis Chen & Zhang 1999 = Oospira jinyungensis;
zhenanensis Chen, Liu & Xu 1996 = Euphaedusa loczyi.

Garnieriinae C. Boettger 1926

Garnieriini

Garnieria Bourguignat 1877
Yunnan.
mouhoti (L. Pfeiffer 1862): Yunnan.

Tropidaucheniini H. Nordsieck 2002

Tropidauchenia Lindholm 1924
Yunnan, Guangxi.
hitomiae H. Nordsieck 2007 (h. hitomiae, h. rufescens H. Nordsieck 2007): Guangxi.
lucida H. Nordsieck 2007 (l. gracillima H. Nordsieck 2007, l. lucida): Guangxi.
nakaharai H. Nordsieck 2007: Guangxi.
napoensis H. Nordsieck 2007: Guangxi.
ootanii H. Nordsieck 2007: Yunnan.
orientalis (Mabille 1887) (o. messageri (Bavay & Dautzenberg 1899), o. orientalis, o. rufocincta H. Nordsieck 2007): Guangxi.

Grandinenia Minato & Chen 1984
Guangxi, Hainan.
amabilis (H. Nordsieck 2005): Guangxi.
fuchsi (Gredler 1883) ( f. cinderella (H. Nordsieck 2002), f. costicollis H. Nordsieck 2007, f. fuchsi, f. pygmaea H. Nordsieck 2007, f. tigris (Chang 2005)): Guangxi.
gastrum (H. Nordsieck 2005) (g. costigera (H. Nordsieck 2005), g. gastrum, g. laticosta (H. Nordsieck 2005), g. pallida H. Nordsieck 2007, g. rubens (H. Nordsieck 2005)): Guangxi.
maroskoi Grego & Szekeres 2011 (m. maroskoi, m. n. subsp. Guangxi): Guangxi.
mirifica (Chen & Gao 1982): Guangxi.
ookuboi (H. Nordsieck 2005) (o. pulchricosta H. Nordsieck 2007, o. ookuboi): Guangxi.
rex H. Nordsieck 2007: Guangxi.
schomburgi (Schmacker & Boettger 1890): Hainan.
takagii (Chang 2004) (t. capreolus (H. Nordsieck 2005), t. gigas H. Nordsieck 2007, t. takagii): Guangxi.
umbra (Chang 2004) (u. pseudofuchsi (H. Nordsieck 2005), u. umbra): Guangxi.

Synonyms

densicostulata H. Nordsieck 2005 = Grandinenia takagii.
kaspari Gredler 1883 = Grandinenia fuchsi;
pusilla Chang 2005 = Grandinenia takagii;
takagii H. Nordsieck 2005 [non Chang 2004] = Grandinenia gastrum.

References

Grego, J. & Szekeres, M. (2011): New taxa of Asiatic Clausiliidae (Mollusca: Gastropoda). – Visaya, 3 (2): 4-22, 4 pls.

International Commission on Zoological Nomenclature (1999): International Code of Zoological Nomenclature. Fourth Edition. – XXIX + 126 pp. (English text).

Nordsieck, H. (2001): Revision of the system of the Phaedusinae from mainland China with the description of new taxa (Gastropoda: Stylommatophora: Clausiliidae). — Arch. Molluskenkunde, 129 (1/2): 25-63, 6 pls.

-- -- -- (2002): Revision of the Garnieriinae (Gastropoda: Stylommatophora: Clausiliidae), with descriptions of new taxa. — Stuttgarter Beitr. Naturkunde (Ser. A), Nr. 640: 23 S..

-- -- -- (2003): Systematic and nomenclatural notes on Phaedusinae with the description of new taxa (Gastropoda: Stylommatophora: Clausiliidae). — Arch. Molluskenkunde, 132 (1/2): 121-141, 3 pls..

-- -- -- (2005): New taxa of Phaedusinae and Garnieriinae from mainland China and Taiwan (Gastropoda: Stylommatophora: Clausiliidae). — Arch. Molluskenkunde, 134 (1): 23-52, 6 pls..

-- -- -- (2007a): Worldwide Door Snails (Clausiliidae), recent and fossil. — 214 pp., 20 pls. Hackenheim (ConchBooks).

-- -- -- (2007b): New taxa of Phaedusinae and Garnieriinae from southern China (Gastropoda: Stylommatophora: Clausiliidae). — Arch. Molluskenkunde, 136 (2): 217-243, 5 pls.

Phaedusinae <

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